Orthogonius

Orthogonius
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Superfamily:	Caraboidea
Family:	Carabidae
Subfamily:	Orthogoniinae
Genus:	Orthogonius; W.S.MacLeay, 1825

Orthogonius is a genus of beetles in the family Carabidae, containing the following species:

Orthogonius aberlenci Tian & Deuve, 2016 (Laos)
Orthogonius acrogonus (Wiedemann, 1819) (Malaysia, Indonesia, and Borneo)
Orthogonius acutangulus Chaudoir, 1878 (Sri Lanka)
Orthogonius adoriae Tian & Deuve, 2006 (India)
Orthogonius aemulus Péringuey, 1896 (Zimbabwe, Botswana, and Namibia)
Orthogonius alternans (Wiedemann, 1823) (Indomalaya)
Orthogonius alutaceus Quedenfeldt, 1883 (Africa)
Orthogonius andrewesi Emden, 1928 (Philippines)
Orthogonius andrewesianus Tian & Deuve, 2006 (Malaysia, Singapore, and Indonesia)
Orthogonius angkor Tian & Deuve, 2006 (Cambodia)
Orthogonius angkorensis Tian & Deuve, 2008 (Cambodia)
Orthogonius angustus Chaudoir, 1872 (Thailand, Vietnam, and Malaysia)
Orthogonius annamensis Jedlicka, 1965 (Vietnam)
Orthogonius annamicola Tian & Deuve, 2006 (Vietnam)
Orthogonius annemariae Tian & Deuve, 2006 (India)
Orthogonius assamensis Tian & Deuve, 2006 (India)
Orthogonius assamicola Tian & Deuve, 2006 (India)
Orthogonius assmuthi Wasmann, 1920 (India)
Orthogonius baconi Chaudoir, 1872 (India and Myanmar)
Orthogonius baconioides Tian & Sabu, 2009 (India)
Orthogonius balthasari Jedlicka, 1935 (Philippines)
Orthogonius batesi Tian & Deuve, 2003 (Sri Lanka)
Orthogonius batesianus Tian & Deuve, 2007
Orthogonius batesioides Tian & Deuve, 2006 (India)
Orthogonius baumi Jedlicka, 1955 (Malaysia)
Orthogonius becqueti Burgeon, 1937 (Democratic Republic of the Congo and Burundi)
Orthogonius bellus Tian & Deuve, 2016 (Vietnam)
Orthogonius bicolor Jedlicka, 1965 (Vietnam)
Orthogonius borneoensis Tian & Deuve, 2006 (Indonesia and Borneo)
Orthogonius brancuccii Tian & Deuve, 2006 (Thailand)
Orthogonius brevilabris Kolbe, 1889 (Democratic Republic of the Congo and Angola)
Orthogonius brevithorax Dejean, 1825 (Africa)
Orthogonius bruggei Tian & Deuve, 2006 (Malaysia)
Orthogonius buquetii Chaudoir, 1850 (Sierra Leone and Ivory Coast)
Orthogonius burckhardti Tian & Deuve, 2006
Orthogonius burmanensis Tian & Deuve, 2006 (Myanmar)
Orthogonius caffer Boheman, 1848 (Zimbabwe and South Africa)
Orthogonius cambodgensis Tian & Deuve, 2006 (Cambodia)
Orthogonius canaraensis Tian & Deuve, 2006 (India)
Orthogonius canaraicola Tian & Deuve, 2006 (India)
Orthogonius capucinus Boheman, 1848 (Zimbabwe and South Africa)
Orthogonius carbonicolor Tian & Deuve, 2006 (Myanmar)
Orthogonius carinatus Tian & Deuve, 2013 (Laos)
Orthogonius celebesicus Tian & Deuve, 2006 (Indonesia)
Orthogonius ceylanicus Tian & Deuve, 2016 (Sri Lanka)
Orthogonius chaudoiri Tian & Deuve, 2001 (Vietnam)
Orthogonius cheni Tian & Deuve, 2001 (China)
Orthogonius chiangensis Tian & Deuve, 2006 (Thailand)
Orthogonius clarkii Murray, 1858 (Africa)
Orthogonius confusus Tian & Deuve, 2006 (Malaysia)
Orthogonius constanti Tian & Deuve, 2006 (Cambodia)
Orthogonius constrictus Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius coomani Tian & Deuve, 2006 (Vietnam)
Orthogonius coomanioides Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius coracinus Kolbe, 1895 (Uganda, Rwanda, Burundi, and Tanzania)
Orthogonius crassicrus Chaudoir, 1872 (Indonesia and Borneo)
Orthogonius cruralis Putzeys in Chaudoir, 1872
Orthogonius curvatus Tian & Deuve, 2006 (India)
Orthogonius cyclothorax Tian & Deuve, 2007 (Singapore)
Orthogonius davidi Chaudoir, 1878 (China and Taiwan)
Orthogonius deletus Schmidt-Goebel, 1846 (China, India, Southeast Asia)
Orthogonius deuvei Tian & Kirschenhofer, 2013 (Vietnam)
Orthogonius dispar Bates, 1892 (Myanmar)
Orthogonius dohertyi Tian & Deuve, 2006 (Malaysia)
Orthogonius dongnanya Tian & Deuve, 2006 (Singapore, Indonesia, and Borneo)
Orthogonius doriae Putzeys in Chaudoir, 1872 (Indonesia and Borneo)
Orthogonius drescheri Liebke, 1937 (Indonesia)
Orthogonius drumonti Tian & Deuve, 2006 (Indonesia)
Orthogonius duboisi Tian & Deuve, 2006 (China)
Orthogonius duplicatus (Wiedemann, 1819) (Indomalaya)
Orthogonius dupuisi Basilewsky, 1948 (Zambia)
Orthogonius dupuisianus Tian & Deuve, 2010 (Indonesia)
Orthogonius dureli Tian & Deuve, 2005 (Bhutan and India)
Orthogonius edentatus Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
Orthogonius elisabethanus Burgeon, 1937 (Democratic Republic of the Congo)
Orthogonius emarginatus Tian & Deuve, 2007 (Singapore)
Orthogonius equimarginalis Tian & Deuve, 2006 (Indonesia)
Orthogonius euthyphallus Tian & Deuve, 2013 (Laos and Vietnam)
Orthogonius fairmairei Tian & Deuve, 2006 (Singapore)
Orthogonius feai Tian & Deuve, 2007
Orthogonius femoralis Chaudoir, 1848 (Sri Lanka and India)
Orthogonius flavipes Deuve, 2004 (Indonesia)
Orthogonius flavus Jedlicka, 1964 (India)
Orthogonius foveiclypeus Tian & Deuve, 2004 (Indonesia)
Orthogonius fryi Tian & Deuve, 2006 (Myanmar)
Orthogonius fugax Chaudoir, 1872 (Sri Lanka)
Orthogonius gracililamella Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius gracilipes Tian & Deuve, 2006 (India)
Orthogonius gracilis Tian & Deuve, 2006 (India)
Orthogonius grootaerti Tian & Deuve, 2006 (Thailand)
Orthogonius hageni Oberthür, 1883 (Indonesia)
Orthogonius himalaya Tian & Deuve, 2006 (Bhutan and India)
Orthogonius himalayicus Tian & Deuve, 2005 (Bhutan and India)
Orthogonius hirtulus Tian & Deuve, 2006 (Indonesia)
Orthogonius hirtus Chaudoir, 1872 (Malaysia and Indonesia)
Orthogonius hopei Gray, 1832 (Indomalaya)
Orthogonius huananoides Tian & Deuve, 2006 (Thailand and Vietnam)
Orthogonius huananus Tian & Deuve, 2001 (China)
Orthogonius hypocrita Chaudoir, 1872 (Indonesia and Philippines)
Orthogonius hypocritoides Jedlicka, 1935 (Philippines)
Orthogonius impunctipennis Quedenfeldt, 1883 (Democratic Republic of the Congo and Angola)
Orthogonius indicus Tian & Deuve, 2006 (India)
Orthogonius indophilus Tian & Deuve, 2006 (India)
Orthogonius inexpectatus Tian & Deuve, 2006 (India)
Orthogonius inops Chaudoir, 1872 (Indonesia)
Orthogonius insularis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
Orthogonius intermedius Chaudoir, 1872 (Indonesia)
Orthogonius javanus Tian & Deuve, 2006 (Taiwan and Indonesia)
Orthogonius jianfengling Tian & Deuve, 2001 (China)
Orthogonius kalimantanensis Tian & Deuve, 2006 (Indonesia and Borneo)
Orthogonius kapiriensis Burgeon, 1937 (Democratic Republic of the Congo)
Orthogonius katangensis Burgeon, 1937 (Democratic Republic of the Congo)
Orthogonius kickeli Kolbe, 1895 (Namibia)
Orthogonius kirirom Tian & Deuve, 2008 (Thailand and Cambodia)
Orthogonius klickai Jedlicka, 1935 (Philippines)
Orthogonius kubani Tian & Deuve, 2006 (Thailand)
Orthogonius kumatai Habu, 1979 (Nepal)
Orthogonius lancangjiang Tian & Deuve, 2006 (Laos)
Orthogonius lao Tian & Deuve, 2006 (Laos)
Orthogonius latreillei Tian & Deuve, 2006 (India)
Orthogonius latus Hope, 1842 (Africa)
Orthogonius ledouxi Tian & Deuve, 2006 (Vietnam)
Orthogonius legrandi Tian & Deuve, 2006 (Malaysia)
Orthogonius lieftincki Andrewes, 1936 (Indonesia)
Orthogonius limbourgi Tian & Deuve, 2016 (Vietnam)
Orthogonius loeicus Tian & Deuve, 2006 (Thailand)
Orthogonius loeiensis Tian & Deuve, 2006 (Thailand)
Orthogonius longicornis Chaudoir, 1872 (Sri Lanka and Thailand)
Orthogonius longilamella Tian & Deuve, 2008
Orthogonius longipennis Hope, 1842
Orthogonius longiphallus Tian & Deuve, 2005 (Bhutan)
Orthogonius lucidus Bates, 1891 (India)
Orthogonius luzonicus Chaudoir, 1872 (Philippines)
Orthogonius macrophthalmus Tian & Deuve, 2013 (Vietnam)
Orthogonius maindronianus Tian & Deuve, 2006 (India)
Orthogonius makiling Tian & Deuve, 2006 (Philippines)
Orthogonius malacca Tian & Deuve, 2006 (Malaysia)
Orthogonius malaisei Andrewes, 1947 (Myanmar)
Orthogonius malaya Tian & Deuve, 2006 (Malaysia)
Orthogonius malaysiaensis Tian & Deuve, 2006 (Malaysia)
Orthogonius medanensis Tian & Deuve, 2006 (Indonesia)
Orthogonius meghalayaensis Tian & Deuve, 2016 (India)
Orthogonius mekongensis Tian & Deuve, 2006 (Laos)
Orthogonius melanipes Tian & Deuve, 2006 (India)
Orthogonius mellyi (Chaudoir, 1850) (Bangladesh, India, and Myanmar)
Orthogonius minimus Tian & Deuve, 2006 (India)
Orthogonius mniszechi Chaudoir, 1872 (India, Malaysia, Indonesia, and Borneo)
Orthogonius moestus Chaudoir, 1872
Orthogonius monolophus Andrewes, 1931 (Indonesia and Borneo)
Orthogonius montanus Tian & Deuve, 2006 (Myanmar)
Orthogonius morvani Tian & Deuve, 2003 (Thailand)
Orthogonius morvanianus Tian & Deuve, 2016 (Thailand)
Orthogonius mouhoti Chaudoir, 1872 (Southeast Asia)
Orthogonius myanmarensis Tian & Deuve, 2006 (Myanmar)
Orthogonius nahaeo Tian & Deuve, 2006 (Thailand)
Orthogonius nepalensis Habu, 1979 (Nepal)
Orthogonius nepalicus Tian & Deuve, 2006 (Nepal)
Orthogonius niger Jedlicka, 1935 (Philippines)
Orthogonius nigripes Tian & Deuve, 2001 (China and Vietnam)
Orthogonius nongfaensis Tian & Kirschenhofer, 2013 (Laos)
Orthogonius nyassicus Kolbe, 1895
Orthogonius obliquepes Tian & Deuve, 2006 (Philippines)
Orthogonius opacus Schmidt-Goebel, 1846 (Indomalaya)
Orthogonius orphnodes Andrewes, 1930 (Indonesia)
Orthogonius ovatulus Tian & Deuve, 2003 (Sri Lanka)
Orthogonius ovatus Chaudoir, 1878 (Indonesia)
Orthogonius pachlatkoi Tian & Deuve, 2006 (Thailand)
Orthogonius palangbangensis Tian & Deuve, 2006 (Indonesia)
Orthogonius panghongae Tian & Deuve, 2006 (China)
Orthogonius pangi Tian & Deuve, 2006 (Thailand and Vietnam)
Orthogonius parallelus Chaudoir, 1872 (Sri Lanka)
Orthogonius parastantoni Tian & Deuve, 2006 (Sri Lanka)
Orthogonius parasuturalis Tian & Deuve, 2006 (Indonesia)
Orthogonius parcepunctatus Kolbe, 1895 (Tanzania)
Orthogonius paris Tian & Deuve, 2006 (Thailand)
Orthogonius parvus Chaudoir, 1872 (India)
Orthogonius pectinatus Tian & Deuve, 2016 (Thailand and Vietnam)
Orthogonius perakensis Tian & Deuve, 2006 (Malaysia)
Orthogonius perakicus Tian & Deuve, 2007 (Malaysia)
Orthogonius perpuncticollis Burgeon, 1937 (Africa)
Orthogonius petiolaris Tian & Deuve, 2006 (India)
Orthogonius piceus Chaudoir, 1872 (Malaysia and Indonesia)
Orthogonius picilabris (W.S.MacLeay, 1825) (Malaysia, Indonesia, and Borneo)
Orthogonius picipennis Chaudoir, 1872 (Cambodia and Vietnam)
Orthogonius pinguis Murray, 1858 (Nigeria)
Orthogonius pinophilus Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius planiger (Walker, 1858) (Sri Lanka)
Orthogonius plebius Tian & Deuve, 2006 (Sri Lanka)
Orthogonius poggii Tian & Deuve, 2007
Orthogonius politior Tian & Deuve, 2016 (Laos)
Orthogonius politus Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
Orthogonius pollinctor Basilewsky, 1948 (Democratic Republic of the Congo)
Orthogonius pradieri Chaudoir, 1872
Orthogonius pseudochaudoiri Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius pseudocheni Tian & Deuve, 2006 (Vietnam)
Orthogonius pseudolongicornis Tian & Deuve, 2006 (Southeast Asia)
Orthogonius pseudomouhoti Tian & Deuve, 2008 (Cambodia)
Orthogonius punctatellus Tian & Deuve, 2006 (Indonesia)
Orthogonius punctulatus Chaudoir, 1872 (India)
Orthogonius punctum Tian & Deuve, 2016 (Thailand and Vietnam)
Orthogonius putzeysi Tian & Deuve, 2006 (Sri Lanka)
Orthogonius quadrisetosus Tian & Deuve, 2006 (Malaysia)
Orthogonius rhodesianus Csiki, 1932
Orthogonius rhodesiensis Lorenz, 1998 (Zambia)
Orthogonius ribbei Tian & Deuve, 2003 (Indonesia)
Orthogonius rogueti Tian & Deuve, 2016 (India)
Orthogonius rotundatus Tian & Deuve, 2006
Orthogonius rufiventris Bates, 1892 (Myanmar)
Orthogonius rufotestaceus G.Müller, 1941 (Ethiopia)
Orthogonius sabahensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
Orthogonius sabahicus Tian & Deuve, 2016 (Malaysia, Indonesia, and Borneo)
Orthogonius salvazai Tian & Deuve, 2006 (Laos)
Orthogonius sarawakensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
Orthogonius saundersi Andrewes, 1926 (Malaysia and Singapore)
Orthogonius schaumi Chaudoir, 1872 (Sri Lanka and Indonesia)
Orthogonius schaumioides Tian & Deuve, 2006 (India)
Orthogonius schauteni Tian & Deuve, 2006 (Malaysia)
Orthogonius schmidtgoebeli Chaudoir, 1872 (Myanmar)
Orthogonius senegalensis Dejean, 1831 (Africa)
Orthogonius setosopalpiger Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius siamensis Tian & Deuve, 2006 (Thailand)
Orthogonius similaris Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius simplicatus Tian & Deuve, 2006 (Indonesia)
Orthogonius singaporensis Tian & Deuve, 2006 (Singapore)
Orthogonius sinuatiphallus Tian & Deuve, 2001 (China)
Orthogonius smetsi Tian & Deuve, 2006 (Cambodia)
Orthogonius solidicornis Tian & Deuve, 2003 (China)
Orthogonius solidus Tian & Deuve, 2006 (India)
Orthogonius spinosus Burgeon, 1937 (Africa)
Orthogonius srilankaicus Tian & Deuve, 2003 (Sri Lanka)
Orthogonius stantoni Tian & Deuve, 2006 (India)
Orthogonius sterbai Jedlicka, 1935 (Philippines)
Orthogonius stygius Andrewes, 1930 (Malaysia and Indonesia)
Orthogonius sulawesicus Tian & Deuve, 2003 (Indonesia)
Orthogonius sulcatoides Tian & Deuve, 2006 (India, Myanmar, and Malaysia)
Orthogonius sulcatus Schmidt-Goebel, 1846 (Myanmar and Cambodia)
Orthogonius sumatraicus Tian & Deuve, 2006 (Indonesia)
Orthogonius suturalis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
Orthogonius taghavianae Tian; Deuve & Felix, 2012 (Thailand)
Orthogonius taiwanicus Tian & Deuve, 2006 (Taiwan)
Orthogonius tangsen Tian & Deuve, 2006 (India)
Orthogonius tenasserimensis Tian & Deuve, 2006 (Myanmar)
Orthogonius termiticola Wasmann, 1902 (Malaysia)
Orthogonius thaicus Tian & Deuve, 2003 (Thailand)
Orthogonius thaiensis Tian & Deuve, 2006 (Thailand)
Orthogonius thailandensis Tian & Deuve, 2006 (Thailand)
Orthogonius thoracicus Gestro, 1875
Orthogonius tikekee Tian & Deuve, 2006 (Myanmar)
Orthogonius tonkinensis Tian & Deuve, 2006 (Vietnam)
Orthogonius tonkinicus Tian & Deuve, 2006 (Vietnam)
Orthogonius travancore Tian & Deuve, 2006 (India)
Orthogonius tricarinatus Burgeon, 1937 (Africa)
Orthogonius uncipennis Tian & Deuve, 2006 (Nepal)
Orthogonius vari Tian; Deuve & Felix, 2012 (Thailand and Cambodia)
Orthogonius variabilis Tian; Deuve & Felix, 2012 (China and Thailand)
Orthogonius virgulatus Andrewes, 1931 (Indonesia and Borneo)
Orthogonius wallardiensis Tian & Deuve, 2006 (India)
Orthogonius wrasei Tian & Deuve, 2016 (Myanmar)
Orthogonius xanthomerus L.Redtenbacher, 1868 (China)
Orthogonius yoga Tian & Deuve, 2006 (India)
Orthogonius yunnanensis Tian & Deuve, 2001 (China)