CRAL-TRIO domain

CRAL/TRIO domain
Alpha-tocopherol transfer protein, closed state with ligand.
Identifiers
SymbolCRAL_TRIO
PfamPF00650
InterProIPR001251
SMARTSec14
SCOP21aua / SCOPe / SUPFAM
OPM superfamily121
OPM protein1r5l
CDDcd00170
Membranome576
Available protein structures:
Pfam  structures / ECOD  
PDBRCSB PDB; PDBe; PDBj
PDBsumstructure summary

CRAL-TRIO domain is a protein structural domain that binds small lipophilic molecules. This domain is named after cellular retinaldehyde-binding protein (CRALBP) and TRIO guanine exchange factor.

CRALB protein carries 11-cis-retinol or 11-cis-retinaldehyde. It modulates interaction of retinoids with visual cycle enzymes. TRIO is involved in coordinating actin remodeling, which is necessary for cell migration and growth.

Other members of the family are alpha-tocopherol transfer protein and phosphatidylinositol-transfer protein (Sec14). They transport their substrates (alpha-tocopherol and phosphatidylinositol or phosphatidylcholine, respectively) between different intracellular membranes. Family also include a guanine nucleotide exchange factor that may function as an effector of RAC1 small G-protein.

The N-terminal domain of yeast ECM25 protein has been identified as containing a lipid binding CRAL-TRIO domain.